Auxilin in enterocytes controls intestinal homeostasis through inter-cell communication

Abstract Residential stem cells sense extrinsic and intrinsic signals to proliferate accordingly to maintain homeostasis. However, how differentiated cells control stem cell proliferation still remains elusive. Here, we find that Auxilin (Aux) maintains enterocyte (EC) integrity to prevent unlimited...

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Main Authors: Runqi Wang, Zhengran Li, Jing Wei, Ruiyan Kong, Xuejing Ren, Hang Zhao, Danjie Zhang, Xiyue Tao Liu, Zhouhua Li
Format: Article
Language:English
Published: Nature Publishing Group 2025-08-01
Series:Cell Death and Disease
Online Access:https://doi.org/10.1038/s41419-025-07954-w
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author Runqi Wang
Zhengran Li
Jing Wei
Ruiyan Kong
Xuejing Ren
Hang Zhao
Danjie Zhang
Xiyue Tao Liu
Zhouhua Li
author_facet Runqi Wang
Zhengran Li
Jing Wei
Ruiyan Kong
Xuejing Ren
Hang Zhao
Danjie Zhang
Xiyue Tao Liu
Zhouhua Li
author_sort Runqi Wang
collection DOAJ
description Abstract Residential stem cells sense extrinsic and intrinsic signals to proliferate accordingly to maintain homeostasis. However, how differentiated cells control stem cell proliferation still remains elusive. Here, we find that Auxilin (Aux) maintains enterocyte (EC) integrity to prevent unlimited intestinal stem cell (ISC) proliferation. Depleting aux in ECs leads to excessive ISC proliferation and intestinal homeostasis disruption. Ectopic cytokine production from dying aux-depleted ECs activates JAK/STAT signaling and promotes ISC proliferation. Mechanistically, Aux facilitates anterograde ER-to-Golgi apparatus (GA) vesicle transport by associating with COPII coatomer. Further, the presentation of cell adhesion molecules (CAMs) by ER-to-GA transport is required for intestinal homeostasis. Together, these data demonstrate that Aux maintains EC integrity by mediating ER-to-GA trafficking of CAMs to restrain excessive ISC proliferation. Thus our study uncovers the underlying mechanism of how differentiated cells control stem cell proliferation through inter-cell communication during tissue homeostasis and pathogenesis.
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institution Kabale University
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publishDate 2025-08-01
publisher Nature Publishing Group
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series Cell Death and Disease
spelling doaj-art-899c36c9c3044c30a04d7ccb32fc1f7e2025-08-24T11:54:27ZengNature Publishing GroupCell Death and Disease2041-48892025-08-0116111310.1038/s41419-025-07954-wAuxilin in enterocytes controls intestinal homeostasis through inter-cell communicationRunqi Wang0Zhengran Li1Jing Wei2Ruiyan Kong3Xuejing Ren4Hang Zhao5Danjie Zhang6Xiyue Tao Liu7Zhouhua Li8Laboratory of Stem Cell Biology, College of Life Sciences, Capital Normal UniversityLaboratory of Stem Cell Biology, College of Life Sciences, Capital Normal UniversityLaboratory of Stem Cell Biology, College of Life Sciences, Capital Normal UniversityLaboratory of Stem Cell Biology, College of Life Sciences, Capital Normal UniversityLaboratory of Stem Cell Biology, College of Life Sciences, Capital Normal UniversityLaboratory of Stem Cell Biology, College of Life Sciences, Capital Normal UniversityLaboratory of Stem Cell Biology, College of Life Sciences, Capital Normal UniversityLaboratory of Stem Cell Biology, College of Life Sciences, Capital Normal UniversityLaboratory of Stem Cell Biology, College of Life Sciences, Capital Normal UniversityAbstract Residential stem cells sense extrinsic and intrinsic signals to proliferate accordingly to maintain homeostasis. However, how differentiated cells control stem cell proliferation still remains elusive. Here, we find that Auxilin (Aux) maintains enterocyte (EC) integrity to prevent unlimited intestinal stem cell (ISC) proliferation. Depleting aux in ECs leads to excessive ISC proliferation and intestinal homeostasis disruption. Ectopic cytokine production from dying aux-depleted ECs activates JAK/STAT signaling and promotes ISC proliferation. Mechanistically, Aux facilitates anterograde ER-to-Golgi apparatus (GA) vesicle transport by associating with COPII coatomer. Further, the presentation of cell adhesion molecules (CAMs) by ER-to-GA transport is required for intestinal homeostasis. Together, these data demonstrate that Aux maintains EC integrity by mediating ER-to-GA trafficking of CAMs to restrain excessive ISC proliferation. Thus our study uncovers the underlying mechanism of how differentiated cells control stem cell proliferation through inter-cell communication during tissue homeostasis and pathogenesis.https://doi.org/10.1038/s41419-025-07954-w
spellingShingle Runqi Wang
Zhengran Li
Jing Wei
Ruiyan Kong
Xuejing Ren
Hang Zhao
Danjie Zhang
Xiyue Tao Liu
Zhouhua Li
Auxilin in enterocytes controls intestinal homeostasis through inter-cell communication
Cell Death and Disease
title Auxilin in enterocytes controls intestinal homeostasis through inter-cell communication
title_full Auxilin in enterocytes controls intestinal homeostasis through inter-cell communication
title_fullStr Auxilin in enterocytes controls intestinal homeostasis through inter-cell communication
title_full_unstemmed Auxilin in enterocytes controls intestinal homeostasis through inter-cell communication
title_short Auxilin in enterocytes controls intestinal homeostasis through inter-cell communication
title_sort auxilin in enterocytes controls intestinal homeostasis through inter cell communication
url https://doi.org/10.1038/s41419-025-07954-w
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