PEG-Mediated Protoplast Transformation of <i>Penicillium sclerotiorum</i> (scaumcx01): Metabolomic Shifts and Root Colonization Dynamics

Protoplast-based transformation is a vital tool for genetic studies in fungi, yet no protoplast method existed for <i>P. sclerotiorum</i>-scaumcx01 before this study. Here, we optimized protoplast isolation, regeneration, and transformation efficiency. The highest protoplast yield (6.72...

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Main Authors: Israt Jahan, Qilin Yang, Zijun Guan, Yihan Wang, Ping Li, Yan Jian
Format: Article
Language:English
Published: MDPI AG 2025-05-01
Series:Journal of Fungi
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Online Access:https://www.mdpi.com/2309-608X/11/5/386
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author Israt Jahan
Qilin Yang
Zijun Guan
Yihan Wang
Ping Li
Yan Jian
author_facet Israt Jahan
Qilin Yang
Zijun Guan
Yihan Wang
Ping Li
Yan Jian
author_sort Israt Jahan
collection DOAJ
description Protoplast-based transformation is a vital tool for genetic studies in fungi, yet no protoplast method existed for <i>P. sclerotiorum</i>-scaumcx01 before this study. Here, we optimized protoplast isolation, regeneration, and transformation efficiency. The highest protoplast yield (6.72 × 10<sup>6</sup> cells/mL) was obtained from liquid mycelium after 12 h of enzymatic digestion at 28 °C using Lysing Enzymes, Yatalase, cellulase, and pectinase. Among osmotic stabilizers, 1 M MgSO<sub>4</sub> yielded the most viable protoplasts. Regeneration occurred via direct mycelial outgrowth and new protoplast formation, with a 1.02% regeneration rate. PEG-mediated transformation with a hygromycin resistance gene and <i>GFP</i> tagging resulted in stable <i>GFP</i> expression in fungal spores and mycelium over five generations. LC/MS-based metabolomic analysis revealed significant changes in glycerophospholipid metabolism, indicating lipid-related dynamics influenced by <i>GFP</i> tagging. Microscopy confirmed successful colonization of tomato roots by <i>GFP</i>-tagged scaumcx01, with <i>GFP</i> fluorescence observed in cortical tissues. Enzymatic (cellulase) seed pretreatment enhanced fungal colonization by modifying root surface properties, promoting plant–fungal interaction. This study establishes an efficient protoplast transformation system, reveals the metabolic impacts of genetic modifications, and demonstrates the potential of enzymatic seed treatment for enhancing plant–fungal interactions.
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publishDate 2025-05-01
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spelling doaj-art-11c9a6fe0bdb4c44a0803d72b711b5c72025-08-20T03:47:54ZengMDPI AGJournal of Fungi2309-608X2025-05-0111538610.3390/jof11050386PEG-Mediated Protoplast Transformation of <i>Penicillium sclerotiorum</i> (scaumcx01): Metabolomic Shifts and Root Colonization DynamicsIsrat Jahan0Qilin Yang1Zijun Guan2Yihan Wang3Ping Li4Yan Jian5Key Laboratory of Agro-Environment in the Tropics, Ministry of Agriculture and Rural Affairs, Guangdong Provincial Key Laboratory of Eco-Circular Agriculture, Guangdong Engineering Research Centre for Modern EcoAgriculture, College of Natural Resources and Environment, South China Agricultural University, Guangzhou 510642, ChinaKey Laboratory of Agro-Environment in the Tropics, Ministry of Agriculture and Rural Affairs, Guangdong Provincial Key Laboratory of Eco-Circular Agriculture, Guangdong Engineering Research Centre for Modern EcoAgriculture, College of Natural Resources and Environment, South China Agricultural University, Guangzhou 510642, ChinaKey Laboratory of Agro-Environment in the Tropics, Ministry of Agriculture and Rural Affairs, Guangdong Provincial Key Laboratory of Eco-Circular Agriculture, Guangdong Engineering Research Centre for Modern EcoAgriculture, College of Natural Resources and Environment, South China Agricultural University, Guangzhou 510642, ChinaKey Laboratory of Agro-Environment in the Tropics, Ministry of Agriculture and Rural Affairs, Guangdong Provincial Key Laboratory of Eco-Circular Agriculture, Guangdong Engineering Research Centre for Modern EcoAgriculture, College of Natural Resources and Environment, South China Agricultural University, Guangzhou 510642, ChinaKey Laboratory of Agro-Environment in the Tropics, Ministry of Agriculture and Rural Affairs, Guangdong Provincial Key Laboratory of Eco-Circular Agriculture, Guangdong Engineering Research Centre for Modern EcoAgriculture, College of Natural Resources and Environment, South China Agricultural University, Guangzhou 510642, ChinaKey Laboratory of Agro-Environment in the Tropics, Ministry of Agriculture and Rural Affairs, Guangdong Provincial Key Laboratory of Eco-Circular Agriculture, Guangdong Engineering Research Centre for Modern EcoAgriculture, College of Natural Resources and Environment, South China Agricultural University, Guangzhou 510642, ChinaProtoplast-based transformation is a vital tool for genetic studies in fungi, yet no protoplast method existed for <i>P. sclerotiorum</i>-scaumcx01 before this study. Here, we optimized protoplast isolation, regeneration, and transformation efficiency. The highest protoplast yield (6.72 × 10<sup>6</sup> cells/mL) was obtained from liquid mycelium after 12 h of enzymatic digestion at 28 °C using Lysing Enzymes, Yatalase, cellulase, and pectinase. Among osmotic stabilizers, 1 M MgSO<sub>4</sub> yielded the most viable protoplasts. Regeneration occurred via direct mycelial outgrowth and new protoplast formation, with a 1.02% regeneration rate. PEG-mediated transformation with a hygromycin resistance gene and <i>GFP</i> tagging resulted in stable <i>GFP</i> expression in fungal spores and mycelium over five generations. LC/MS-based metabolomic analysis revealed significant changes in glycerophospholipid metabolism, indicating lipid-related dynamics influenced by <i>GFP</i> tagging. Microscopy confirmed successful colonization of tomato roots by <i>GFP</i>-tagged scaumcx01, with <i>GFP</i> fluorescence observed in cortical tissues. Enzymatic (cellulase) seed pretreatment enhanced fungal colonization by modifying root surface properties, promoting plant–fungal interaction. This study establishes an efficient protoplast transformation system, reveals the metabolic impacts of genetic modifications, and demonstrates the potential of enzymatic seed treatment for enhancing plant–fungal interactions.https://www.mdpi.com/2309-608X/11/5/386<i>Penicillium sclerotiorum</i>protoplast transformation<i>GFP</i> taggingmetabolomic analysisplant–fungal interaction
spellingShingle Israt Jahan
Qilin Yang
Zijun Guan
Yihan Wang
Ping Li
Yan Jian
PEG-Mediated Protoplast Transformation of <i>Penicillium sclerotiorum</i> (scaumcx01): Metabolomic Shifts and Root Colonization Dynamics
Journal of Fungi
<i>Penicillium sclerotiorum</i>
protoplast transformation
<i>GFP</i> tagging
metabolomic analysis
plant–fungal interaction
title PEG-Mediated Protoplast Transformation of <i>Penicillium sclerotiorum</i> (scaumcx01): Metabolomic Shifts and Root Colonization Dynamics
title_full PEG-Mediated Protoplast Transformation of <i>Penicillium sclerotiorum</i> (scaumcx01): Metabolomic Shifts and Root Colonization Dynamics
title_fullStr PEG-Mediated Protoplast Transformation of <i>Penicillium sclerotiorum</i> (scaumcx01): Metabolomic Shifts and Root Colonization Dynamics
title_full_unstemmed PEG-Mediated Protoplast Transformation of <i>Penicillium sclerotiorum</i> (scaumcx01): Metabolomic Shifts and Root Colonization Dynamics
title_short PEG-Mediated Protoplast Transformation of <i>Penicillium sclerotiorum</i> (scaumcx01): Metabolomic Shifts and Root Colonization Dynamics
title_sort peg mediated protoplast transformation of i penicillium sclerotiorum i scaumcx01 metabolomic shifts and root colonization dynamics
topic <i>Penicillium sclerotiorum</i>
protoplast transformation
<i>GFP</i> tagging
metabolomic analysis
plant–fungal interaction
url https://www.mdpi.com/2309-608X/11/5/386
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