Bmal1 haploinsufficiency impairs fear memory and modulates neuroinflammation via the 5-HT2C receptor

BackgroundBMAL1, a key regulator of circadian rhythms, plays a multifaceted role in brain function. However, the complex interplay between BMAL1, memory, neuroinflammation, and neurotransmitter regulation remains poorly understood. To investigate these interactions, we conducted a study using BMAL1-...

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Main Authors: Weifen Li, Shengnan Mou, Tahir Ali, Tianxiang Li, Yan Liu, Shupeng Li, Xiaoming Yu, Zhi-Jian Yu
Format: Article
Language:English
Published: Frontiers Media S.A. 2024-11-01
Series:Frontiers in Pharmacology
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Online Access:https://www.frontiersin.org/articles/10.3389/fphar.2024.1422693/full
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author Weifen Li
Weifen Li
Shengnan Mou
Tahir Ali
Tahir Ali
Tianxiang Li
Yan Liu
Shupeng Li
Shupeng Li
Shupeng Li
Xiaoming Yu
Zhi-Jian Yu
author_facet Weifen Li
Weifen Li
Shengnan Mou
Tahir Ali
Tahir Ali
Tianxiang Li
Yan Liu
Shupeng Li
Shupeng Li
Shupeng Li
Xiaoming Yu
Zhi-Jian Yu
author_sort Weifen Li
collection DOAJ
description BackgroundBMAL1, a key regulator of circadian rhythms, plays a multifaceted role in brain function. However, the complex interplay between BMAL1, memory, neuroinflammation, and neurotransmitter regulation remains poorly understood. To investigate these interactions, we conducted a study using BMAL1-haplodeficient mice (BMAL1+/−).MethodsWe exposed BMAL1+/− mice to behavioral assessments including cued fear conditioning, new objection recognition (NOR) test, and Y-maze test to evaluate BMAL1+/− haplodeficiency impact on memory. Furthermore, biochemical changes were analyzed through western blotting, and ELISA to explore further the mechanism of BMAL1+/− in memory, and neuroinflammation.ResultsWe found that BMAL1 haploinsufficiency led to deficits in cued fear learning and memory, while spatial memory and object recognition remained intact. Further analysis revealed dysregulated neurotransmitter levels and alterations in neurotransmitter-related proteins in the prefrontal cortex of BMAL1+/− mice. Pharmacological interventions targeting dopamine uptake or the 5-HT2C receptor demonstrated that inhibiting the 5-HT2C receptor could rescue fear learning and memory impairments in BMAL1+/− mice. Additionally, we observed downregulation of the inflammasome and neuroinflammation pathways in BMAL1+/− mice, which is validated by inflammation mediator lipopolysaccharide (LPS) administration.ConclusionThese findings highlight that BMAL1 haploinsufficiency leads to deficits in fear learning and memory, which are linked to alterations in neurotransmitters and receptors, particularly the 5-HT2C receptor. Targeting the 5-HT2C receptor may offer a potential therapeutic strategy for mitigating cognitive impairments associated with BMAL1 dysfunction.
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spelling doaj-art-f0ab59b4e7394795a3a593bcaa25a0652024-11-14T04:47:47ZengFrontiers Media S.A.Frontiers in Pharmacology1663-98122024-11-011510.3389/fphar.2024.14226931422693Bmal1 haploinsufficiency impairs fear memory and modulates neuroinflammation via the 5-HT2C receptorWeifen Li0Weifen Li1Shengnan Mou2Tahir Ali3Tahir Ali4Tianxiang Li5Yan Liu6Shupeng Li7Shupeng Li8Shupeng Li9Xiaoming Yu10Zhi-Jian Yu11School of Pharmacy, Shenzhen University Medical School, Shenzhen University, Shenzhen, ChinaDepartment of Infectious Diseases and Shenzhen Key Laboratory for Endogenous Infections, The 6th Affiliated Hospital of Shenzhen University Health Science Center, Shenzhen, ChinaState Key Laboratory of Oncogenomics, School of Chemical Biology and Biotechnology, Peking University Shenzhen Graduate School, Shenzhen, ChinaState Key Laboratory of Oncogenomics, School of Chemical Biology and Biotechnology, Peking University Shenzhen Graduate School, Shenzhen, ChinaShenzhen Bay Laboratory, Shenzhen, ChinaState Key Laboratory of Oncogenomics, School of Chemical Biology and Biotechnology, Peking University Shenzhen Graduate School, Shenzhen, ChinaThe Seventh Affiliated Hospital of Sun Yat-Sen University, Shenzhen, ChinaState Key Laboratory of Oncogenomics, School of Chemical Biology and Biotechnology, Peking University Shenzhen Graduate School, Shenzhen, ChinaShenzhen Bay Laboratory, Shenzhen, ChinaDepartment of Psychiatry, University of Toronto, Toronto, ON, CanadaCancer Center, The Second Hospital, Cheeloo College of Medicine, Shandong University, Jinan, ChinaDepartment of Infectious Diseases and Shenzhen Key Laboratory for Endogenous Infections, The 6th Affiliated Hospital of Shenzhen University Health Science Center, Shenzhen, ChinaBackgroundBMAL1, a key regulator of circadian rhythms, plays a multifaceted role in brain function. However, the complex interplay between BMAL1, memory, neuroinflammation, and neurotransmitter regulation remains poorly understood. To investigate these interactions, we conducted a study using BMAL1-haplodeficient mice (BMAL1+/−).MethodsWe exposed BMAL1+/− mice to behavioral assessments including cued fear conditioning, new objection recognition (NOR) test, and Y-maze test to evaluate BMAL1+/− haplodeficiency impact on memory. Furthermore, biochemical changes were analyzed through western blotting, and ELISA to explore further the mechanism of BMAL1+/− in memory, and neuroinflammation.ResultsWe found that BMAL1 haploinsufficiency led to deficits in cued fear learning and memory, while spatial memory and object recognition remained intact. Further analysis revealed dysregulated neurotransmitter levels and alterations in neurotransmitter-related proteins in the prefrontal cortex of BMAL1+/− mice. Pharmacological interventions targeting dopamine uptake or the 5-HT2C receptor demonstrated that inhibiting the 5-HT2C receptor could rescue fear learning and memory impairments in BMAL1+/− mice. Additionally, we observed downregulation of the inflammasome and neuroinflammation pathways in BMAL1+/− mice, which is validated by inflammation mediator lipopolysaccharide (LPS) administration.ConclusionThese findings highlight that BMAL1 haploinsufficiency leads to deficits in fear learning and memory, which are linked to alterations in neurotransmitters and receptors, particularly the 5-HT2C receptor. Targeting the 5-HT2C receptor may offer a potential therapeutic strategy for mitigating cognitive impairments associated with BMAL1 dysfunction.https://www.frontiersin.org/articles/10.3389/fphar.2024.1422693/fullBmal1learning and memory defects5-HT2CRneuroinflammationneurotransmitter
spellingShingle Weifen Li
Weifen Li
Shengnan Mou
Tahir Ali
Tahir Ali
Tianxiang Li
Yan Liu
Shupeng Li
Shupeng Li
Shupeng Li
Xiaoming Yu
Zhi-Jian Yu
Bmal1 haploinsufficiency impairs fear memory and modulates neuroinflammation via the 5-HT2C receptor
Frontiers in Pharmacology
Bmal1
learning and memory defects
5-HT2CR
neuroinflammation
neurotransmitter
title Bmal1 haploinsufficiency impairs fear memory and modulates neuroinflammation via the 5-HT2C receptor
title_full Bmal1 haploinsufficiency impairs fear memory and modulates neuroinflammation via the 5-HT2C receptor
title_fullStr Bmal1 haploinsufficiency impairs fear memory and modulates neuroinflammation via the 5-HT2C receptor
title_full_unstemmed Bmal1 haploinsufficiency impairs fear memory and modulates neuroinflammation via the 5-HT2C receptor
title_short Bmal1 haploinsufficiency impairs fear memory and modulates neuroinflammation via the 5-HT2C receptor
title_sort bmal1 haploinsufficiency impairs fear memory and modulates neuroinflammation via the 5 ht2c receptor
topic Bmal1
learning and memory defects
5-HT2CR
neuroinflammation
neurotransmitter
url https://www.frontiersin.org/articles/10.3389/fphar.2024.1422693/full
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