Case Report: A novel CXCR4 variant (p.S341Y) in a family with a pathogenic NFKB1 variant and variable clinical manifestations

WHIM syndrome is typically caused by C-terminal gain-of-function variants in CXCR4, yet clinical heterogeneity suggests additional genetic modifiers. We investigated a family in which the 22-year-old proband harbored two heterozygous variants: a novel CXCR4 missense variant, c.1022C>A (p.S341...

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Main Authors: Melis Yilmaz, Katarina Zmajkovicova, Rahim Z. Miller, Grace Blair, Maryssa Ellison, Boglarka Ujhazi, Maria Chitty Lopez, Joseph F. Dasso, Jacob R. Bledsoe, Krisztian Csomos, Barbara Maierhofer, Adriana Badarau, Joao P. Pereira, Henry Kanarek, Christoph B. Geier, Jolan E. Walter
Format: Article
Language:English
Published: Frontiers Media S.A. 2025-08-01
Series:Frontiers in Immunology
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Online Access:https://www.frontiersin.org/articles/10.3389/fimmu.2025.1641122/full
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author Melis Yilmaz
Katarina Zmajkovicova
Rahim Z. Miller
Rahim Z. Miller
Grace Blair
Maryssa Ellison
Boglarka Ujhazi
Maria Chitty Lopez
Maria Chitty Lopez
Joseph F. Dasso
Jacob R. Bledsoe
Krisztian Csomos
Barbara Maierhofer
Adriana Badarau
Joao P. Pereira
Henry Kanarek
Christoph B. Geier
Christoph B. Geier
Jolan E. Walter
Jolan E. Walter
author_facet Melis Yilmaz
Katarina Zmajkovicova
Rahim Z. Miller
Rahim Z. Miller
Grace Blair
Maryssa Ellison
Boglarka Ujhazi
Maria Chitty Lopez
Maria Chitty Lopez
Joseph F. Dasso
Jacob R. Bledsoe
Krisztian Csomos
Barbara Maierhofer
Adriana Badarau
Joao P. Pereira
Henry Kanarek
Christoph B. Geier
Christoph B. Geier
Jolan E. Walter
Jolan E. Walter
author_sort Melis Yilmaz
collection DOAJ
description WHIM syndrome is typically caused by C-terminal gain-of-function variants in CXCR4, yet clinical heterogeneity suggests additional genetic modifiers. We investigated a family in which the 22-year-old proband harbored two heterozygous variants: a novel CXCR4 missense variant, c.1022C>A (p.S341Y), and a frameshift variant in NFKB1, c.980dup (p.A328Sfs*12). Functionally, CXCR4 p.S341Y substitution - located two residues upstream of the known pathogenic p.E343K variant - increased CXCL12-induced chemotaxis and ERK/AKT signaling while minimally affecting receptor internalization, supporting a partial CXCR4 gain-of-function. The CXCR4 variant co-segregated with mild neutropenia, recurrent respiratory infections, and cutaneous warts in the paternal lineage. In contrast, the maternal NFKB1 variant was associated with agammaglobulinemia and autoimmunity. Their co-inheritance in the proband resulted in a blended WHIM/CVID phenotype characterized by myelokathexis, B-cell maturation arrest and T-cell dysregulation. This case expands the phenotypic spectrum of CXCR4 variants and highlights how multilocus inheritance can obscure classical diagnostic boundaries and guide individualized therapy.
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spelling doaj-art-c7ea4d428a404fef890bcbe8b385f26c2025-08-20T05:32:31ZengFrontiers Media S.A.Frontiers in Immunology1664-32242025-08-011610.3389/fimmu.2025.16411221641122Case Report: A novel CXCR4 variant (p.S341Y) in a family with a pathogenic NFKB1 variant and variable clinical manifestationsMelis Yilmaz0Katarina Zmajkovicova1Rahim Z. Miller2Rahim Z. Miller3Grace Blair4Maryssa Ellison5Boglarka Ujhazi6Maria Chitty Lopez7Maria Chitty Lopez8Joseph F. Dasso9Jacob R. Bledsoe10Krisztian Csomos11Barbara Maierhofer12Adriana Badarau13Joao P. Pereira14Henry Kanarek15Christoph B. Geier16Christoph B. Geier17Jolan E. Walter18Jolan E. Walter19Division of Allergy and Immunology, Department of Pediatrics and Medicine, Morsani College of Medicine, University of South Florida at Johns Hopkins All Children’s Hospital, St. Petersburg, FL, United StatesResearch Department, Formerly X4 Pharmaceuticals (Austria) GmbH, Vienna, AustriaDivision of Allergy and Immunology, Department of Pediatrics and Medicine, Morsani College of Medicine, University of South Florida at Johns Hopkins All Children’s Hospital, St. Petersburg, FL, United StatesDivision of Allergy and Immunology, Department of Pediatrics, Johns Hopkins All Children’s Hospital, St. Petersburg, FL, United StatesDivision of Allergy and Immunology, Department of Pediatrics and Medicine, Morsani College of Medicine, University of South Florida at Johns Hopkins All Children’s Hospital, St. Petersburg, FL, United StatesDivision of Allergy and Immunology, Department of Pediatrics and Medicine, Morsani College of Medicine, University of South Florida at Johns Hopkins All Children’s Hospital, St. Petersburg, FL, United StatesDivision of Allergy and Immunology, Department of Pediatrics and Medicine, Morsani College of Medicine, University of South Florida at Johns Hopkins All Children’s Hospital, St. Petersburg, FL, United StatesDivision of Allergy and Immunology, Department of Pediatrics and Medicine, Morsani College of Medicine, University of South Florida at Johns Hopkins All Children’s Hospital, St. Petersburg, FL, United StatesDivision of Allergy and Immunology, Department of Pediatrics, Johns Hopkins All Children’s Hospital, St. Petersburg, FL, United StatesDivision of Allergy and Immunology, Department of Pediatrics and Medicine, Morsani College of Medicine, University of South Florida at Johns Hopkins All Children’s Hospital, St. Petersburg, FL, United StatesDepartment of Pathology, Boston Children’s Hospital, Harvard Medical School, Boston, MA, United StatesDivision of Allergy and Immunology, Department of Pediatrics and Medicine, Morsani College of Medicine, University of South Florida at Johns Hopkins All Children’s Hospital, St. Petersburg, FL, United StatesResearch Department, Formerly X4 Pharmaceuticals (Austria) GmbH, Vienna, AustriaResearch Department, Formerly X4 Pharmaceuticals (Austria) GmbH, Vienna, AustriaDepartment of Immunobiology, Yale School of Medicine, Yale University, New Haven, CT, United StatesKanarek Allergy, Asthma & Immunology, Overland Park, KS, United StatesDivision of Immunology, Faculty of Medicine and Health Sciences, University Medicine Oldenburg, Oldenburg, GermanyInstitute of Medical Genetics, Faculty of Medicine and Health Sciences, University Medicine Oldenburg, Oldenburg, GermanyDivision of Allergy and Immunology, Department of Pediatrics and Medicine, Morsani College of Medicine, University of South Florida at Johns Hopkins All Children’s Hospital, St. Petersburg, FL, United StatesDivision of Allergy and Immunology, Department of Pediatrics, Johns Hopkins All Children’s Hospital, St. Petersburg, FL, United StatesWHIM syndrome is typically caused by C-terminal gain-of-function variants in CXCR4, yet clinical heterogeneity suggests additional genetic modifiers. We investigated a family in which the 22-year-old proband harbored two heterozygous variants: a novel CXCR4 missense variant, c.1022C>A (p.S341Y), and a frameshift variant in NFKB1, c.980dup (p.A328Sfs*12). Functionally, CXCR4 p.S341Y substitution - located two residues upstream of the known pathogenic p.E343K variant - increased CXCL12-induced chemotaxis and ERK/AKT signaling while minimally affecting receptor internalization, supporting a partial CXCR4 gain-of-function. The CXCR4 variant co-segregated with mild neutropenia, recurrent respiratory infections, and cutaneous warts in the paternal lineage. In contrast, the maternal NFKB1 variant was associated with agammaglobulinemia and autoimmunity. Their co-inheritance in the proband resulted in a blended WHIM/CVID phenotype characterized by myelokathexis, B-cell maturation arrest and T-cell dysregulation. This case expands the phenotypic spectrum of CXCR4 variants and highlights how multilocus inheritance can obscure classical diagnostic boundaries and guide individualized therapy.https://www.frontiersin.org/articles/10.3389/fimmu.2025.1641122/fullCXCR4WHIMNFKB1neutropeniaCVID
spellingShingle Melis Yilmaz
Katarina Zmajkovicova
Rahim Z. Miller
Rahim Z. Miller
Grace Blair
Maryssa Ellison
Boglarka Ujhazi
Maria Chitty Lopez
Maria Chitty Lopez
Joseph F. Dasso
Jacob R. Bledsoe
Krisztian Csomos
Barbara Maierhofer
Adriana Badarau
Joao P. Pereira
Henry Kanarek
Christoph B. Geier
Christoph B. Geier
Jolan E. Walter
Jolan E. Walter
Case Report: A novel CXCR4 variant (p.S341Y) in a family with a pathogenic NFKB1 variant and variable clinical manifestations
Frontiers in Immunology
CXCR4
WHIM
NFKB1
neutropenia
CVID
title Case Report: A novel CXCR4 variant (p.S341Y) in a family with a pathogenic NFKB1 variant and variable clinical manifestations
title_full Case Report: A novel CXCR4 variant (p.S341Y) in a family with a pathogenic NFKB1 variant and variable clinical manifestations
title_fullStr Case Report: A novel CXCR4 variant (p.S341Y) in a family with a pathogenic NFKB1 variant and variable clinical manifestations
title_full_unstemmed Case Report: A novel CXCR4 variant (p.S341Y) in a family with a pathogenic NFKB1 variant and variable clinical manifestations
title_short Case Report: A novel CXCR4 variant (p.S341Y) in a family with a pathogenic NFKB1 variant and variable clinical manifestations
title_sort case report a novel cxcr4 variant p s341y in a family with a pathogenic nfkb1 variant and variable clinical manifestations
topic CXCR4
WHIM
NFKB1
neutropenia
CVID
url https://www.frontiersin.org/articles/10.3389/fimmu.2025.1641122/full
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