Plasma membrane remodeling determines adipocyte expansion and mechanical adaptability

Abstract Adipocytes expand massively to accommodate excess energy stores and protect the organism from lipotoxicity. Adipose tissue expandability is at the center of disorders such as obesity and lipodystrophy; however, little is known about the relevance of adipocyte biomechanics on the etiology of...

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Main Authors: María C. M. Aboy-Pardal, Marta C. Guadamillas, Carlos R. Guerrero, Mauro Català-Montoro, Mónica Toledano-Donado, Sara Terrés-Domínguez, Dácil M. Pavón, Víctor Jiménez-Jiménez, Daniel Jimenez-Carretero, Moreno Zamai, Cintia Folgueira, Ana Cerezo, Fidel-Nicolás Lolo, Rubén Nogueiras, Guadalupe Sabio, Miguel Sánchez-Álvarez, Asier Echarri, Ricardo Garcia, Miguel A. Del Pozo
Format: Article
Language:English
Published: Nature Portfolio 2024-11-01
Series:Nature Communications
Online Access:https://doi.org/10.1038/s41467-024-54224-y
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author María C. M. Aboy-Pardal
Marta C. Guadamillas
Carlos R. Guerrero
Mauro Català-Montoro
Mónica Toledano-Donado
Sara Terrés-Domínguez
Dácil M. Pavón
Víctor Jiménez-Jiménez
Daniel Jimenez-Carretero
Moreno Zamai
Cintia Folgueira
Ana Cerezo
Fidel-Nicolás Lolo
Rubén Nogueiras
Guadalupe Sabio
Miguel Sánchez-Álvarez
Asier Echarri
Ricardo Garcia
Miguel A. Del Pozo
author_facet María C. M. Aboy-Pardal
Marta C. Guadamillas
Carlos R. Guerrero
Mauro Català-Montoro
Mónica Toledano-Donado
Sara Terrés-Domínguez
Dácil M. Pavón
Víctor Jiménez-Jiménez
Daniel Jimenez-Carretero
Moreno Zamai
Cintia Folgueira
Ana Cerezo
Fidel-Nicolás Lolo
Rubén Nogueiras
Guadalupe Sabio
Miguel Sánchez-Álvarez
Asier Echarri
Ricardo Garcia
Miguel A. Del Pozo
author_sort María C. M. Aboy-Pardal
collection DOAJ
description Abstract Adipocytes expand massively to accommodate excess energy stores and protect the organism from lipotoxicity. Adipose tissue expandability is at the center of disorders such as obesity and lipodystrophy; however, little is known about the relevance of adipocyte biomechanics on the etiology of these conditions. Here, we show in male mice in vivo that the adipocyte plasma membrane undergoes caveolar domain reorganization upon lipid droplet expansion. As the lipid droplet grows, caveolae disassemble to release their membrane reservoir and increase cell surface area, and transfer specific caveolar components to the LD surface. Adipose tissue null for caveolae is stiffer, shows compromised deformability, and is prone to rupture under mechanical compression. Mechanistically, phosphoacceptor Cav1 Tyr14 is required for caveolae disassembly: adipocytes bearing a Tyr14Phe mutation at this residue are stiffer and smaller, leading to decreased adiposity in vivo; exhibit deficient transfer of Cav1 and EHD2 to the LD surface, and show distinct Cav1 molecular dynamics and tension adaptation. These results indicate that Cav1 phosphoregulation modulates caveolar dynamics as a relevant component of the homeostatic mechanoadaptation of the differentiated adipocyte.
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spelling doaj-art-67c1dc358d36404a815842f71018ebb32024-12-01T12:35:43ZengNature PortfolioNature Communications2041-17232024-11-0115112710.1038/s41467-024-54224-yPlasma membrane remodeling determines adipocyte expansion and mechanical adaptabilityMaría C. M. Aboy-Pardal0Marta C. Guadamillas1Carlos R. Guerrero2Mauro Català-Montoro3Mónica Toledano-Donado4Sara Terrés-Domínguez5Dácil M. Pavón6Víctor Jiménez-Jiménez7Daniel Jimenez-Carretero8Moreno Zamai9Cintia Folgueira10Ana Cerezo11Fidel-Nicolás Lolo12Rubén Nogueiras13Guadalupe Sabio14Miguel Sánchez-Álvarez15Asier Echarri16Ricardo Garcia17Miguel A. Del Pozo18Mechanoadaptation and Caveolae Biology lab, Novel mechanisms in atherosclerosis program. Centro Nacional de Investigaciones Cardiovasculares (CNIC)Mechanoadaptation and Caveolae Biology lab, Novel mechanisms in atherosclerosis program. Centro Nacional de Investigaciones Cardiovasculares (CNIC)ForceTool group, Instituto de Ciencia de Materiales de Madrid (ICMM), CSICMechanoadaptation and Caveolae Biology lab, Novel mechanisms in atherosclerosis program. Centro Nacional de Investigaciones Cardiovasculares (CNIC)Mechanoadaptation and Caveolae Biology lab, Novel mechanisms in atherosclerosis program. Centro Nacional de Investigaciones Cardiovasculares (CNIC)Mechanoadaptation and Caveolae Biology lab, Novel mechanisms in atherosclerosis program. Centro Nacional de Investigaciones Cardiovasculares (CNIC)Mechanoadaptation and Caveolae Biology lab, Novel mechanisms in atherosclerosis program. Centro Nacional de Investigaciones Cardiovasculares (CNIC)Mechanoadaptation and Caveolae Biology lab, Novel mechanisms in atherosclerosis program. Centro Nacional de Investigaciones Cardiovasculares (CNIC)Cellomics Unit, Cell and Developmental Biology Area. Centro Nacional de Investigaciones Cardiovasculares (CNIC)Microscopy and Dynamic Imaging Unit, Centro Nacional de Investigaciones Cardiovasculares (CNIC)Centro de Investigación en Medicina Molecular y Enfermedades Crónicas CIMUSMechanoadaptation and Caveolae Biology lab, Novel mechanisms in atherosclerosis program. Centro Nacional de Investigaciones Cardiovasculares (CNIC)Mechanoadaptation and Caveolae Biology lab, Novel mechanisms in atherosclerosis program. Centro Nacional de Investigaciones Cardiovasculares (CNIC)Centro de Investigación en Medicina Molecular y Enfermedades Crónicas CIMUSStress kinases in Diabetes, Cancer and Cardiovascular Disease lab. Cardiovascular risk factors & brain function program, Centro Nacional de Investigaciones Cardiovasculares (CNIC)Mechanoadaptation and Caveolae Biology lab, Novel mechanisms in atherosclerosis program. Centro Nacional de Investigaciones Cardiovasculares (CNIC)Mechanoadaptation and Caveolae Biology lab, Novel mechanisms in atherosclerosis program. Centro Nacional de Investigaciones Cardiovasculares (CNIC)ForceTool group, Instituto de Ciencia de Materiales de Madrid (ICMM), CSICMechanoadaptation and Caveolae Biology lab, Novel mechanisms in atherosclerosis program. Centro Nacional de Investigaciones Cardiovasculares (CNIC)Abstract Adipocytes expand massively to accommodate excess energy stores and protect the organism from lipotoxicity. Adipose tissue expandability is at the center of disorders such as obesity and lipodystrophy; however, little is known about the relevance of adipocyte biomechanics on the etiology of these conditions. Here, we show in male mice in vivo that the adipocyte plasma membrane undergoes caveolar domain reorganization upon lipid droplet expansion. As the lipid droplet grows, caveolae disassemble to release their membrane reservoir and increase cell surface area, and transfer specific caveolar components to the LD surface. Adipose tissue null for caveolae is stiffer, shows compromised deformability, and is prone to rupture under mechanical compression. Mechanistically, phosphoacceptor Cav1 Tyr14 is required for caveolae disassembly: adipocytes bearing a Tyr14Phe mutation at this residue are stiffer and smaller, leading to decreased adiposity in vivo; exhibit deficient transfer of Cav1 and EHD2 to the LD surface, and show distinct Cav1 molecular dynamics and tension adaptation. These results indicate that Cav1 phosphoregulation modulates caveolar dynamics as a relevant component of the homeostatic mechanoadaptation of the differentiated adipocyte.https://doi.org/10.1038/s41467-024-54224-y
spellingShingle María C. M. Aboy-Pardal
Marta C. Guadamillas
Carlos R. Guerrero
Mauro Català-Montoro
Mónica Toledano-Donado
Sara Terrés-Domínguez
Dácil M. Pavón
Víctor Jiménez-Jiménez
Daniel Jimenez-Carretero
Moreno Zamai
Cintia Folgueira
Ana Cerezo
Fidel-Nicolás Lolo
Rubén Nogueiras
Guadalupe Sabio
Miguel Sánchez-Álvarez
Asier Echarri
Ricardo Garcia
Miguel A. Del Pozo
Plasma membrane remodeling determines adipocyte expansion and mechanical adaptability
Nature Communications
title Plasma membrane remodeling determines adipocyte expansion and mechanical adaptability
title_full Plasma membrane remodeling determines adipocyte expansion and mechanical adaptability
title_fullStr Plasma membrane remodeling determines adipocyte expansion and mechanical adaptability
title_full_unstemmed Plasma membrane remodeling determines adipocyte expansion and mechanical adaptability
title_short Plasma membrane remodeling determines adipocyte expansion and mechanical adaptability
title_sort plasma membrane remodeling determines adipocyte expansion and mechanical adaptability
url https://doi.org/10.1038/s41467-024-54224-y
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