The miR-6240 target gene Igf2bp3 promotes myoblast fusion by enhancing myomaker mRNA stability

Abstract Background Myoblast fusion plays a crucial role in myogenesis. Insulin-like growth factor 2 mRNA-binding protein 3 (IGF2BP3) functions as an RNA N 6-methyladenosine reader and exerts important roles in various biological processes. While our prior study suggested Igf2bp3 contributes to myog...

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Main Authors: Yuxin Huang, Wei Wang, Xinhao Fan, Xiaoqin Liu, Weiwei Liu, Zishuai Wang, Yixing Li, Yalan Yang, Zhonglin Tang
Format: Article
Language:English
Published: BMC 2024-12-01
Series:Cellular & Molecular Biology Letters
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Online Access:https://doi.org/10.1186/s11658-024-00650-1
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author Yuxin Huang
Wei Wang
Xinhao Fan
Xiaoqin Liu
Weiwei Liu
Zishuai Wang
Yixing Li
Yalan Yang
Zhonglin Tang
author_facet Yuxin Huang
Wei Wang
Xinhao Fan
Xiaoqin Liu
Weiwei Liu
Zishuai Wang
Yixing Li
Yalan Yang
Zhonglin Tang
author_sort Yuxin Huang
collection DOAJ
description Abstract Background Myoblast fusion plays a crucial role in myogenesis. Insulin-like growth factor 2 mRNA-binding protein 3 (IGF2BP3) functions as an RNA N 6-methyladenosine reader and exerts important roles in various biological processes. While our prior study suggested Igf2bp3 contributes to myogenesis, its molecular regulatory mechanism is largely unclear. Methods Real-time quantitative polymerase chain reaction (RT-qPCR) and western blot were used for gene expression analysis. siRNA and CRISPRi technologies were conducted to knockdown the expression of Igf2bp3. CRISPR/Cas9 technology was performed to knockout Igf2bp3. The Igf2bp3 overexpression vector was designed using the pcDNA3.1(+) vector. Immunofluorescence detection was employed for subcellular localization and cell differentiation analysis. Cell Counting Kit-8 (CCK-8) and 5-ethynyl-2′-deoxyuridine (EdU) assays were conducted for cell proliferation and fusion detection. The dual-luciferase reporter assay and RNA immunoprecipitation (RIP) assay were utilized for regulatory mechanism analysis of Igf2bp3. Results The overexpression of Igf2bp3 enhances myoblast fusion while knockdown of Igf2bp3 blocks the formation of myotubes. miR-6240 promotes myoblast proliferation while preventing myoblast differentiation and fusion by targeting the 3′ untranslated rgion (UTR) of Igf2bp3. Notably, the impacts of miR-6240 mimics on myoblast proliferation, differentiation, and fusion can be effectively counteracted by the overexpression of Igf2bp3. Moreover, our findings elucidate a direct interaction between Igf2bp3 and the myoblast fusion factor myomaker (Mymk). Igf2bp3 binds to Mymk to enhance its mRNA stability. This interaction results in increased expression of Mymk and heightened myoblast fusion. Conclusions Our study unveils Igf2bp3 as a novel post-transcriptional regulator of myoblast fusion through the miR-6240/Mymk axis, significantly contributing to our understanding of skeletal muscle development. Graphical Abstract
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institution Kabale University
issn 1689-1392
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series Cellular & Molecular Biology Letters
spelling doaj-art-4b6c8ab6bcca49cf8090d4bd6ab3195e2024-12-08T12:37:24ZengBMCCellular & Molecular Biology Letters1689-13922024-12-0129112010.1186/s11658-024-00650-1The miR-6240 target gene Igf2bp3 promotes myoblast fusion by enhancing myomaker mRNA stabilityYuxin Huang0Wei Wang1Xinhao Fan2Xiaoqin Liu3Weiwei Liu4Zishuai Wang5Yixing Li6Yalan Yang7Zhonglin Tang8Kunpeng Institute of Modern Agriculture at Foshan, Agricultural Genomics Institute at Shenzhen, Chinese Academy of Agricultural SciencesKunpeng Institute of Modern Agriculture at Foshan, Agricultural Genomics Institute at Shenzhen, Chinese Academy of Agricultural SciencesKunpeng Institute of Modern Agriculture at Foshan, Agricultural Genomics Institute at Shenzhen, Chinese Academy of Agricultural SciencesKunpeng Institute of Modern Agriculture at Foshan, Agricultural Genomics Institute at Shenzhen, Chinese Academy of Agricultural SciencesKunpeng Institute of Modern Agriculture at Foshan, Agricultural Genomics Institute at Shenzhen, Chinese Academy of Agricultural SciencesKunpeng Institute of Modern Agriculture at Foshan, Agricultural Genomics Institute at Shenzhen, Chinese Academy of Agricultural SciencesGuangxi Key Laboratory of Animal Breeding, Disease Control and Prevention; College of Animal Science and Technology, Guangxi UniversityKunpeng Institute of Modern Agriculture at Foshan, Agricultural Genomics Institute at Shenzhen, Chinese Academy of Agricultural SciencesKunpeng Institute of Modern Agriculture at Foshan, Agricultural Genomics Institute at Shenzhen, Chinese Academy of Agricultural SciencesAbstract Background Myoblast fusion plays a crucial role in myogenesis. Insulin-like growth factor 2 mRNA-binding protein 3 (IGF2BP3) functions as an RNA N 6-methyladenosine reader and exerts important roles in various biological processes. While our prior study suggested Igf2bp3 contributes to myogenesis, its molecular regulatory mechanism is largely unclear. Methods Real-time quantitative polymerase chain reaction (RT-qPCR) and western blot were used for gene expression analysis. siRNA and CRISPRi technologies were conducted to knockdown the expression of Igf2bp3. CRISPR/Cas9 technology was performed to knockout Igf2bp3. The Igf2bp3 overexpression vector was designed using the pcDNA3.1(+) vector. Immunofluorescence detection was employed for subcellular localization and cell differentiation analysis. Cell Counting Kit-8 (CCK-8) and 5-ethynyl-2′-deoxyuridine (EdU) assays were conducted for cell proliferation and fusion detection. The dual-luciferase reporter assay and RNA immunoprecipitation (RIP) assay were utilized for regulatory mechanism analysis of Igf2bp3. Results The overexpression of Igf2bp3 enhances myoblast fusion while knockdown of Igf2bp3 blocks the formation of myotubes. miR-6240 promotes myoblast proliferation while preventing myoblast differentiation and fusion by targeting the 3′ untranslated rgion (UTR) of Igf2bp3. Notably, the impacts of miR-6240 mimics on myoblast proliferation, differentiation, and fusion can be effectively counteracted by the overexpression of Igf2bp3. Moreover, our findings elucidate a direct interaction between Igf2bp3 and the myoblast fusion factor myomaker (Mymk). Igf2bp3 binds to Mymk to enhance its mRNA stability. This interaction results in increased expression of Mymk and heightened myoblast fusion. Conclusions Our study unveils Igf2bp3 as a novel post-transcriptional regulator of myoblast fusion through the miR-6240/Mymk axis, significantly contributing to our understanding of skeletal muscle development. Graphical Abstracthttps://doi.org/10.1186/s11658-024-00650-1Igf2bp3Myobalst fusionMymkmRNA stabilitymiR-6240Myogenesis
spellingShingle Yuxin Huang
Wei Wang
Xinhao Fan
Xiaoqin Liu
Weiwei Liu
Zishuai Wang
Yixing Li
Yalan Yang
Zhonglin Tang
The miR-6240 target gene Igf2bp3 promotes myoblast fusion by enhancing myomaker mRNA stability
Cellular & Molecular Biology Letters
Igf2bp3
Myobalst fusion
Mymk
mRNA stability
miR-6240
Myogenesis
title The miR-6240 target gene Igf2bp3 promotes myoblast fusion by enhancing myomaker mRNA stability
title_full The miR-6240 target gene Igf2bp3 promotes myoblast fusion by enhancing myomaker mRNA stability
title_fullStr The miR-6240 target gene Igf2bp3 promotes myoblast fusion by enhancing myomaker mRNA stability
title_full_unstemmed The miR-6240 target gene Igf2bp3 promotes myoblast fusion by enhancing myomaker mRNA stability
title_short The miR-6240 target gene Igf2bp3 promotes myoblast fusion by enhancing myomaker mRNA stability
title_sort mir 6240 target gene igf2bp3 promotes myoblast fusion by enhancing myomaker mrna stability
topic Igf2bp3
Myobalst fusion
Mymk
mRNA stability
miR-6240
Myogenesis
url https://doi.org/10.1186/s11658-024-00650-1
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