HADHA Regulates Respiratory Complex Assembly and Couples FAO and OXPHOS

Abstract Oxidative phosphorylation (OXPHOS) and fatty acid oxidation (FAO) are key bioenergetics pathways. The machineries for both processes are localized in mitochondria. Secondary OXPHOS defects have been documented in patients with primary FAO deficiencies, and vice versa. However, the underlyin...

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Main Authors: Chaoying Qin, Shasha Gong, Ting Liang, Zhenbo Zhang, Jessie Thomas, Janice Deng, Yaguang Liu, Peiqing Hu, Bi Zhu, Shujie Song, Marisol Fernández Ortiz, Yuji Ikeno, Exing Wang, James Lechleiter, Susan T. Weintraub, Yidong Bai
Format: Article
Language:English
Published: Wiley 2024-12-01
Series:Advanced Science
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Online Access:https://doi.org/10.1002/advs.202405147
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author Chaoying Qin
Shasha Gong
Ting Liang
Zhenbo Zhang
Jessie Thomas
Janice Deng
Yaguang Liu
Peiqing Hu
Bi Zhu
Shujie Song
Marisol Fernández Ortiz
Yuji Ikeno
Exing Wang
James Lechleiter
Susan T. Weintraub
Yidong Bai
author_facet Chaoying Qin
Shasha Gong
Ting Liang
Zhenbo Zhang
Jessie Thomas
Janice Deng
Yaguang Liu
Peiqing Hu
Bi Zhu
Shujie Song
Marisol Fernández Ortiz
Yuji Ikeno
Exing Wang
James Lechleiter
Susan T. Weintraub
Yidong Bai
author_sort Chaoying Qin
collection DOAJ
description Abstract Oxidative phosphorylation (OXPHOS) and fatty acid oxidation (FAO) are key bioenergetics pathways. The machineries for both processes are localized in mitochondria. Secondary OXPHOS defects have been documented in patients with primary FAO deficiencies, and vice versa. However, the underlying mechanisms remain unclear. Intrigued by the observations that regulation of supercomplexes (SCs) assembly in a mouse OXPHOS deficient cell line and its derivatives is associated with the changes in lipid metabolism, a proteomics analysis is carried out and identified mitochondrial trifunctional protein (MTP) subunit alpha (hydroxyacyl‐CoA dehydrogenase trifunctional multienzyme complex subunit alpha, HADHA) as a potential regulatory factor for SCs assembly. HADHA‐Knockdown cells and mouse embryonic fibroblasts (MEFs) derived from HADHA‐Knockout mice displayed both reduced SCs assembly and defective OXPHOS. Stimulation of OXPHOS induced in cell culture by replacing glucose with galactose and of lipid metabolism in mice with a high‐fat diet (HFD) both exhibited increased HADHA expression. HADHA Heterozygous mice fed with HFD showed enhanced steatosis associated with a reduction of SCs assembly and OXPHOS function. The results indicate that HADHA participates in SCs assembly and couples FAO and OXPHOS.
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spelling doaj-art-4a65a6e132004df783d3baf65f858ef12024-12-18T14:18:10ZengWileyAdvanced Science2198-38442024-12-011147n/an/a10.1002/advs.202405147HADHA Regulates Respiratory Complex Assembly and Couples FAO and OXPHOSChaoying Qin0Shasha Gong1Ting Liang2Zhenbo Zhang3Jessie Thomas4Janice Deng5Yaguang Liu6Peiqing Hu7Bi Zhu8Shujie Song9Marisol Fernández Ortiz10Yuji Ikeno11Exing Wang12James Lechleiter13Susan T. Weintraub14Yidong Bai15Department of Cell Systems and Anatomy The University of Texas Health San Antonio San AntonioTexas 78229 USADepartment of Cell Systems and Anatomy The University of Texas Health San Antonio San AntonioTexas 78229 USADepartment of Cell Systems and Anatomy The University of Texas Health San Antonio San AntonioTexas 78229 USADepartment of Cell Systems and Anatomy The University of Texas Health San Antonio San AntonioTexas 78229 USADepartment of Cell Systems and Anatomy The University of Texas Health San Antonio San AntonioTexas 78229 USADepartment of Cell Systems and Anatomy The University of Texas Health San Antonio San AntonioTexas 78229 USADepartment of Cell Systems and Anatomy The University of Texas Health San Antonio San AntonioTexas 78229 USADepartment of Cell Systems and Anatomy The University of Texas Health San Antonio San AntonioTexas 78229 USADepartment of Cell Systems and Anatomy The University of Texas Health San Antonio San AntonioTexas 78229 USADepartment of Cell Systems and Anatomy The University of Texas Health San Antonio San AntonioTexas 78229 USADepartment of Cell Systems and Anatomy The University of Texas Health San Antonio San AntonioTexas 78229 USABarshop Institute of Aging Research and Longevity and Department of Pathology University of Texas Health San Antonio San AntonioTexas 78229 USADepartment of Cell Systems and Anatomy The University of Texas Health San Antonio San AntonioTexas 78229 USADepartment of Cell Systems and Anatomy The University of Texas Health San Antonio San AntonioTexas 78229 USADepartment of Biochemistry and Structural Biology The University of Texas Health San Antonio San AntonioTexas 78229 USADepartment of Cell Systems and Anatomy The University of Texas Health San Antonio San AntonioTexas 78229 USAAbstract Oxidative phosphorylation (OXPHOS) and fatty acid oxidation (FAO) are key bioenergetics pathways. The machineries for both processes are localized in mitochondria. Secondary OXPHOS defects have been documented in patients with primary FAO deficiencies, and vice versa. However, the underlying mechanisms remain unclear. Intrigued by the observations that regulation of supercomplexes (SCs) assembly in a mouse OXPHOS deficient cell line and its derivatives is associated with the changes in lipid metabolism, a proteomics analysis is carried out and identified mitochondrial trifunctional protein (MTP) subunit alpha (hydroxyacyl‐CoA dehydrogenase trifunctional multienzyme complex subunit alpha, HADHA) as a potential regulatory factor for SCs assembly. HADHA‐Knockdown cells and mouse embryonic fibroblasts (MEFs) derived from HADHA‐Knockout mice displayed both reduced SCs assembly and defective OXPHOS. Stimulation of OXPHOS induced in cell culture by replacing glucose with galactose and of lipid metabolism in mice with a high‐fat diet (HFD) both exhibited increased HADHA expression. HADHA Heterozygous mice fed with HFD showed enhanced steatosis associated with a reduction of SCs assembly and OXPHOS function. The results indicate that HADHA participates in SCs assembly and couples FAO and OXPHOS.https://doi.org/10.1002/advs.202405147fatty acid oxidation (FAO)HADHAmitochondrial respiratory chainmitochondrial trifunctional protein (MTP)respiratory complex I
spellingShingle Chaoying Qin
Shasha Gong
Ting Liang
Zhenbo Zhang
Jessie Thomas
Janice Deng
Yaguang Liu
Peiqing Hu
Bi Zhu
Shujie Song
Marisol Fernández Ortiz
Yuji Ikeno
Exing Wang
James Lechleiter
Susan T. Weintraub
Yidong Bai
HADHA Regulates Respiratory Complex Assembly and Couples FAO and OXPHOS
Advanced Science
fatty acid oxidation (FAO)
HADHA
mitochondrial respiratory chain
mitochondrial trifunctional protein (MTP)
respiratory complex I
title HADHA Regulates Respiratory Complex Assembly and Couples FAO and OXPHOS
title_full HADHA Regulates Respiratory Complex Assembly and Couples FAO and OXPHOS
title_fullStr HADHA Regulates Respiratory Complex Assembly and Couples FAO and OXPHOS
title_full_unstemmed HADHA Regulates Respiratory Complex Assembly and Couples FAO and OXPHOS
title_short HADHA Regulates Respiratory Complex Assembly and Couples FAO and OXPHOS
title_sort hadha regulates respiratory complex assembly and couples fao and oxphos
topic fatty acid oxidation (FAO)
HADHA
mitochondrial respiratory chain
mitochondrial trifunctional protein (MTP)
respiratory complex I
url https://doi.org/10.1002/advs.202405147
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