HURP regulates Kif18A recruitment and activity to synergistically control microtubule dynamics

Abstract During mitosis, microtubule dynamics are regulated to ensure proper alignment and segregation of chromosomes. The dynamics of kinetochore-attached microtubules are regulated by hepatoma-upregulated protein (HURP) and the mitotic kinesin-8 Kif18A, but the underlying mechanism remains elusive...

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Main Authors: Juan M. Perez-Bertoldi, Yuanchang Zhao, Akanksha Thawani, Ahmet Yildiz, Eva Nogales
Format: Article
Language:English
Published: Nature Portfolio 2024-11-01
Series:Nature Communications
Online Access:https://doi.org/10.1038/s41467-024-53691-7
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author Juan M. Perez-Bertoldi
Yuanchang Zhao
Akanksha Thawani
Ahmet Yildiz
Eva Nogales
author_facet Juan M. Perez-Bertoldi
Yuanchang Zhao
Akanksha Thawani
Ahmet Yildiz
Eva Nogales
author_sort Juan M. Perez-Bertoldi
collection DOAJ
description Abstract During mitosis, microtubule dynamics are regulated to ensure proper alignment and segregation of chromosomes. The dynamics of kinetochore-attached microtubules are regulated by hepatoma-upregulated protein (HURP) and the mitotic kinesin-8 Kif18A, but the underlying mechanism remains elusive. Using single-molecule imaging in vitro, we demonstrate that Kif18A motility is regulated by HURP. While sparse decoration of HURP activates the motor, higher concentrations hinder processive motility. To shed light on this behavior, we determine the binding mode of HURP to microtubules using cryo-EM. The structure helps rationalize why HURP functions as a microtubule stabilizer. Additionally, HURP partially overlaps with the microtubule-binding site of the Kif18A motor domain, indicating that excess HURP inhibits Kif18A motility by steric exclusion. We also observe that HURP and Kif18A function together to suppress dynamics of the microtubule plus-end, providing a mechanistic basis for how they collectively serve in microtubule length control.
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institution Kabale University
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spelling doaj-art-3d658b7fa25c4e8486b0237bfceba6572024-11-10T12:31:34ZengNature PortfolioNature Communications2041-17232024-11-0115111810.1038/s41467-024-53691-7HURP regulates Kif18A recruitment and activity to synergistically control microtubule dynamicsJuan M. Perez-Bertoldi0Yuanchang Zhao1Akanksha Thawani2Ahmet Yildiz3Eva Nogales4Biophysics Graduate Group, University of CaliforniaPhysics Department, University of CaliforniaDepartment of Molecular and Cell Biology, University of CaliforniaBiophysics Graduate Group, University of CaliforniaDepartment of Molecular and Cell Biology, University of CaliforniaAbstract During mitosis, microtubule dynamics are regulated to ensure proper alignment and segregation of chromosomes. The dynamics of kinetochore-attached microtubules are regulated by hepatoma-upregulated protein (HURP) and the mitotic kinesin-8 Kif18A, but the underlying mechanism remains elusive. Using single-molecule imaging in vitro, we demonstrate that Kif18A motility is regulated by HURP. While sparse decoration of HURP activates the motor, higher concentrations hinder processive motility. To shed light on this behavior, we determine the binding mode of HURP to microtubules using cryo-EM. The structure helps rationalize why HURP functions as a microtubule stabilizer. Additionally, HURP partially overlaps with the microtubule-binding site of the Kif18A motor domain, indicating that excess HURP inhibits Kif18A motility by steric exclusion. We also observe that HURP and Kif18A function together to suppress dynamics of the microtubule plus-end, providing a mechanistic basis for how they collectively serve in microtubule length control.https://doi.org/10.1038/s41467-024-53691-7
spellingShingle Juan M. Perez-Bertoldi
Yuanchang Zhao
Akanksha Thawani
Ahmet Yildiz
Eva Nogales
HURP regulates Kif18A recruitment and activity to synergistically control microtubule dynamics
Nature Communications
title HURP regulates Kif18A recruitment and activity to synergistically control microtubule dynamics
title_full HURP regulates Kif18A recruitment and activity to synergistically control microtubule dynamics
title_fullStr HURP regulates Kif18A recruitment and activity to synergistically control microtubule dynamics
title_full_unstemmed HURP regulates Kif18A recruitment and activity to synergistically control microtubule dynamics
title_short HURP regulates Kif18A recruitment and activity to synergistically control microtubule dynamics
title_sort hurp regulates kif18a recruitment and activity to synergistically control microtubule dynamics
url https://doi.org/10.1038/s41467-024-53691-7
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