Seipin governs phosphatidic acid homeostasis at the inner nuclear membrane

Abstract The nuclear envelope is a specialized subdomain of the endoplasmic reticulum and comprises the inner and outer nuclear membranes. Despite the crucial role of the inner nuclear membrane in genome regulation, its lipid metabolism remains poorly understood. Phosphatidic acid (PA) is essential...

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Main Authors: Anete Romanauska, Edvinas Stankunas, Maya Schuldiner, Alwin Köhler
Format: Article
Language:English
Published: Nature Portfolio 2024-12-01
Series:Nature Communications
Online Access:https://doi.org/10.1038/s41467-024-54811-z
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author Anete Romanauska
Edvinas Stankunas
Maya Schuldiner
Alwin Köhler
author_facet Anete Romanauska
Edvinas Stankunas
Maya Schuldiner
Alwin Köhler
author_sort Anete Romanauska
collection DOAJ
description Abstract The nuclear envelope is a specialized subdomain of the endoplasmic reticulum and comprises the inner and outer nuclear membranes. Despite the crucial role of the inner nuclear membrane in genome regulation, its lipid metabolism remains poorly understood. Phosphatidic acid (PA) is essential for membrane growth as well as lipid storage. Using a genome-wide lipid biosensor screen in S. cerevisiae, we identify regulators of inner nuclear membrane PA homeostasis, including yeast Seipin, a known mediator of nuclear lipid droplet biogenesis. Here, we show that Seipin preserves nuclear envelope integrity by preventing its deformation and ectopic membrane formation. Mutations of specific regions of Seipin, some linked to human lipodystrophy, disrupt PA distribution at the inner nuclear membrane and nuclear lipid droplet formation. Investigating the Seipin co-factor Ldb16 reveals that a triacylglycerol binding site is crucial for lipid droplet formation, whereas PA regulation can be functionally separated. Our study highlights the potential of lipid biosensor screens for examining inner nuclear membrane lipid metabolism.
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institution Kabale University
issn 2041-1723
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publishDate 2024-12-01
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spelling doaj-art-33a363b2bf4d4162a5b440976abae7c62024-12-08T12:36:09ZengNature PortfolioNature Communications2041-17232024-12-0115111610.1038/s41467-024-54811-zSeipin governs phosphatidic acid homeostasis at the inner nuclear membraneAnete Romanauska0Edvinas Stankunas1Maya Schuldiner2Alwin Köhler3Max Perutz Labs, Vienna BioCenter Campus (VBC), Dr. Bohr-Gasse 9/3Max Perutz Labs, Vienna BioCenter Campus (VBC), Dr. Bohr-Gasse 9/3Department of Molecular Genetics, Weizmann Institute of ScienceMax Perutz Labs, Vienna BioCenter Campus (VBC), Dr. Bohr-Gasse 9/3Abstract The nuclear envelope is a specialized subdomain of the endoplasmic reticulum and comprises the inner and outer nuclear membranes. Despite the crucial role of the inner nuclear membrane in genome regulation, its lipid metabolism remains poorly understood. Phosphatidic acid (PA) is essential for membrane growth as well as lipid storage. Using a genome-wide lipid biosensor screen in S. cerevisiae, we identify regulators of inner nuclear membrane PA homeostasis, including yeast Seipin, a known mediator of nuclear lipid droplet biogenesis. Here, we show that Seipin preserves nuclear envelope integrity by preventing its deformation and ectopic membrane formation. Mutations of specific regions of Seipin, some linked to human lipodystrophy, disrupt PA distribution at the inner nuclear membrane and nuclear lipid droplet formation. Investigating the Seipin co-factor Ldb16 reveals that a triacylglycerol binding site is crucial for lipid droplet formation, whereas PA regulation can be functionally separated. Our study highlights the potential of lipid biosensor screens for examining inner nuclear membrane lipid metabolism.https://doi.org/10.1038/s41467-024-54811-z
spellingShingle Anete Romanauska
Edvinas Stankunas
Maya Schuldiner
Alwin Köhler
Seipin governs phosphatidic acid homeostasis at the inner nuclear membrane
Nature Communications
title Seipin governs phosphatidic acid homeostasis at the inner nuclear membrane
title_full Seipin governs phosphatidic acid homeostasis at the inner nuclear membrane
title_fullStr Seipin governs phosphatidic acid homeostasis at the inner nuclear membrane
title_full_unstemmed Seipin governs phosphatidic acid homeostasis at the inner nuclear membrane
title_short Seipin governs phosphatidic acid homeostasis at the inner nuclear membrane
title_sort seipin governs phosphatidic acid homeostasis at the inner nuclear membrane
url https://doi.org/10.1038/s41467-024-54811-z
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