Nanocarrier-mediated RNAi of CYP9A306 and CYB5R enhances susceptibility of invasive tomato pest, Tuta absoluta to cyantraniliprole

Cyantraniliprole, a second-generation anthranilic diamide, is widely used to manage lepidopteran pests, including the invasive tomato pinworm Tuta absoluta (Meyrick). However, cyantraniliprole’s resistance mechanisms and associated fitness costs in T. absoluta remain underexplored. Here, we investig...

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Main Authors: Farman Ullah, Guru-Pirasanna-Pandi G, Hina Gul, Rudra Madhab Panda, Ghulam Murtaza, Zhijun Zhang, Jun Huang, Xiaowei Li, Nicolas Desneux, Yaobin Lu
Format: Article
Language:English
Published: Frontiers Media S.A. 2025-04-01
Series:Frontiers in Plant Science
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Online Access:https://www.frontiersin.org/articles/10.3389/fpls.2025.1573634/full
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author Farman Ullah
Guru-Pirasanna-Pandi G
Hina Gul
Rudra Madhab Panda
Ghulam Murtaza
Zhijun Zhang
Jun Huang
Xiaowei Li
Nicolas Desneux
Yaobin Lu
author_facet Farman Ullah
Guru-Pirasanna-Pandi G
Hina Gul
Rudra Madhab Panda
Ghulam Murtaza
Zhijun Zhang
Jun Huang
Xiaowei Li
Nicolas Desneux
Yaobin Lu
author_sort Farman Ullah
collection DOAJ
description Cyantraniliprole, a second-generation anthranilic diamide, is widely used to manage lepidopteran pests, including the invasive tomato pinworm Tuta absoluta (Meyrick). However, cyantraniliprole’s resistance mechanisms and associated fitness costs in T. absoluta remain underexplored. Here, we investigated the fitness costs and resistance mechanisms of cyantraniliprole-resistant strain (CyanRS) via nanocarrier-mediated RNA interference (RNAi). Results showed that the egg incubation period and developmental durations of larval and pupal stages were significantly prolonged in the CyanRS population of T. absoluta compared to the susceptible strain (SS). Further, the adult emergence, longevities of male and female, fecundity, and hatching rate were significantly reduced in CyanRS individuals. The mRNA expression levels of cytochrome b5 reductase (CYB5R) and cytochrome P450 (CYP9A306) were analyzed using RT-qPCR to explore their potential involvement in cyantraniliprole resistance in T. absoluta. Phylogenetic and motif analysis of CYB5R and CYP9A306 indicated their evolutionary and functional conservation with other insect species, especially Lepidopterans. Notably, nanocarrier-encapsulated dsRNA targeting CYB5R and CYP9A306 genes significantly reduced their expression levels. Further, the activity of cytochrome P450 was substantially decreased after the knockdown of the CYB5R and CYP9A306 genes. This increased susceptibility of the resistant population of T. absoluta to cyantraniliprole, leading to a higher mortality rate than the controls. These findings show that CYB5R and CYP9A306 might play a key role in cyantraniliprole resistance evolution in T. absoluta. The current study provides in-depth insights to understand the underlying mechanisms of cyantraniliprole resistance in this key invasive herbivore.
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spelling doaj-art-09d4af0d4cf24ce8bf8cbb7d0a2d58b92025-08-20T03:29:23ZengFrontiers Media S.A.Frontiers in Plant Science1664-462X2025-04-011610.3389/fpls.2025.15736341573634Nanocarrier-mediated RNAi of CYP9A306 and CYB5R enhances susceptibility of invasive tomato pest, Tuta absoluta to cyantraniliproleFarman Ullah0Guru-Pirasanna-Pandi G1Hina Gul2Rudra Madhab Panda3Ghulam Murtaza4Zhijun Zhang5Jun Huang6Xiaowei Li7Nicolas Desneux8Yaobin Lu9State Key Laboratory for Quality and Safety of Agro-Products, Key Laboratory of Biotechnology in Plant Protection of MOA of China and Zhejiang Province, Institute of Plant Protection and Microbiology, Zhejiang Academy of Agricultural Sciences, Hangzhou, ChinaProtection Division, ICAR-National Rice Research Institute, Cuttack, Odisha, IndiaState Key Laboratory for Quality and Safety of Agro-Products, Key Laboratory of Biotechnology in Plant Protection of MOA of China and Zhejiang Province, Institute of Plant Protection and Microbiology, Zhejiang Academy of Agricultural Sciences, Hangzhou, ChinaProtection Division, ICAR-National Rice Research Institute, Cuttack, Odisha, IndiaMARA Key Laboratory of Surveillance and Management for Plant Quarantine Pests, College of Plant Protection, China Agricultural University, Beijing, ChinaState Key Laboratory for Quality and Safety of Agro-Products, Key Laboratory of Biotechnology in Plant Protection of MOA of China and Zhejiang Province, Institute of Plant Protection and Microbiology, Zhejiang Academy of Agricultural Sciences, Hangzhou, ChinaState Key Laboratory for Quality and Safety of Agro-Products, Key Laboratory of Biotechnology in Plant Protection of MOA of China and Zhejiang Province, Institute of Plant Protection and Microbiology, Zhejiang Academy of Agricultural Sciences, Hangzhou, ChinaState Key Laboratory for Quality and Safety of Agro-Products, Key Laboratory of Biotechnology in Plant Protection of MOA of China and Zhejiang Province, Institute of Plant Protection and Microbiology, Zhejiang Academy of Agricultural Sciences, Hangzhou, ChinaUniversité Côte d’Azur, INRAE, CNRS, UMR ISA, Nice, FranceState Key Laboratory for Quality and Safety of Agro-Products, Key Laboratory of Biotechnology in Plant Protection of MOA of China and Zhejiang Province, Institute of Plant Protection and Microbiology, Zhejiang Academy of Agricultural Sciences, Hangzhou, ChinaCyantraniliprole, a second-generation anthranilic diamide, is widely used to manage lepidopteran pests, including the invasive tomato pinworm Tuta absoluta (Meyrick). However, cyantraniliprole’s resistance mechanisms and associated fitness costs in T. absoluta remain underexplored. Here, we investigated the fitness costs and resistance mechanisms of cyantraniliprole-resistant strain (CyanRS) via nanocarrier-mediated RNA interference (RNAi). Results showed that the egg incubation period and developmental durations of larval and pupal stages were significantly prolonged in the CyanRS population of T. absoluta compared to the susceptible strain (SS). Further, the adult emergence, longevities of male and female, fecundity, and hatching rate were significantly reduced in CyanRS individuals. The mRNA expression levels of cytochrome b5 reductase (CYB5R) and cytochrome P450 (CYP9A306) were analyzed using RT-qPCR to explore their potential involvement in cyantraniliprole resistance in T. absoluta. Phylogenetic and motif analysis of CYB5R and CYP9A306 indicated their evolutionary and functional conservation with other insect species, especially Lepidopterans. Notably, nanocarrier-encapsulated dsRNA targeting CYB5R and CYP9A306 genes significantly reduced their expression levels. Further, the activity of cytochrome P450 was substantially decreased after the knockdown of the CYB5R and CYP9A306 genes. This increased susceptibility of the resistant population of T. absoluta to cyantraniliprole, leading to a higher mortality rate than the controls. These findings show that CYB5R and CYP9A306 might play a key role in cyantraniliprole resistance evolution in T. absoluta. The current study provides in-depth insights to understand the underlying mechanisms of cyantraniliprole resistance in this key invasive herbivore.https://www.frontiersin.org/articles/10.3389/fpls.2025.1573634/fullresistance evolutionRNA interferencebiological traitslepidopteragene expression
spellingShingle Farman Ullah
Guru-Pirasanna-Pandi G
Hina Gul
Rudra Madhab Panda
Ghulam Murtaza
Zhijun Zhang
Jun Huang
Xiaowei Li
Nicolas Desneux
Yaobin Lu
Nanocarrier-mediated RNAi of CYP9A306 and CYB5R enhances susceptibility of invasive tomato pest, Tuta absoluta to cyantraniliprole
Frontiers in Plant Science
resistance evolution
RNA interference
biological traits
lepidoptera
gene expression
title Nanocarrier-mediated RNAi of CYP9A306 and CYB5R enhances susceptibility of invasive tomato pest, Tuta absoluta to cyantraniliprole
title_full Nanocarrier-mediated RNAi of CYP9A306 and CYB5R enhances susceptibility of invasive tomato pest, Tuta absoluta to cyantraniliprole
title_fullStr Nanocarrier-mediated RNAi of CYP9A306 and CYB5R enhances susceptibility of invasive tomato pest, Tuta absoluta to cyantraniliprole
title_full_unstemmed Nanocarrier-mediated RNAi of CYP9A306 and CYB5R enhances susceptibility of invasive tomato pest, Tuta absoluta to cyantraniliprole
title_short Nanocarrier-mediated RNAi of CYP9A306 and CYB5R enhances susceptibility of invasive tomato pest, Tuta absoluta to cyantraniliprole
title_sort nanocarrier mediated rnai of cyp9a306 and cyb5r enhances susceptibility of invasive tomato pest tuta absoluta to cyantraniliprole
topic resistance evolution
RNA interference
biological traits
lepidoptera
gene expression
url https://www.frontiersin.org/articles/10.3389/fpls.2025.1573634/full
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