Enhancing drought tolerance in faba bean using ascorbic and humic acids: role of antioxidant enzymes and compatible solutes

Abstract Drought is a major environmental stress, particularly in arid regions, where it severely limits faba bean productivity. Foliar-applied ascorbic acid (AsA) and soil-applied humic acid (HA) significantly improved drought tolerance in three faba bean cultivars by enhancing physiological perfor...

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Main Authors: Hussein S. Al-Demrdash, Anam Ayyoub, Omar E. A. ziton, Saber A.E. Mowafy, El-Sayed E. A. El-Sayed, Uthman Balgith Algopishi, Ahmed Ezzat Ahmed, Khaled A. El-Tarabily, Synan F. AbuQamar, Mohsin Mahmood, El-Sayed M. Desoky
Format: Article
Language:English
Published: BMC 2025-08-01
Series:BMC Plant Biology
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Online Access:https://doi.org/10.1186/s12870-025-06971-3
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author Hussein S. Al-Demrdash
Anam Ayyoub
Omar E. A. ziton
Saber A.E. Mowafy
El-Sayed E. A. El-Sayed
Uthman Balgith Algopishi
Ahmed Ezzat Ahmed
Khaled A. El-Tarabily
Synan F. AbuQamar
Mohsin Mahmood
El-Sayed M. Desoky
author_facet Hussein S. Al-Demrdash
Anam Ayyoub
Omar E. A. ziton
Saber A.E. Mowafy
El-Sayed E. A. El-Sayed
Uthman Balgith Algopishi
Ahmed Ezzat Ahmed
Khaled A. El-Tarabily
Synan F. AbuQamar
Mohsin Mahmood
El-Sayed M. Desoky
author_sort Hussein S. Al-Demrdash
collection DOAJ
description Abstract Drought is a major environmental stress, particularly in arid regions, where it severely limits faba bean productivity. Foliar-applied ascorbic acid (AsA) and soil-applied humic acid (HA) significantly improved drought tolerance in three faba bean cultivars by enhancing physiological performance and mitigating oxidative damage under moderate (300 mm) and severe (200 mm) drought conditions. Drought stress significantly reduced chlorophyll content (up to -57.5%), relative water content (RWC, -37.9%), and yield traits such as plant height (− 9.6%) and seed yield (− 20.8%), while increasing oxidative stress markers like malondialdehyde (MDA, + 192.8%) and hydrogen peroxide (H₂O₂, + 105.0%). AsA and HA alleviated these effects, improving chlorophyll retention (up to + 33.7%), water status (+ 17.0%), and reducing MDA(− 19.1%) and electrolyte leakage (− 11.5%). Enhanced accumulation of proline (+ 27.4%) and soluble sugars (+ 18.0%) contributed to improved osmotic balance, while antioxidant enzyme activities (superoxide dismutase, catalase, peroxidase, ascorbate peroxidase, and glutathione reductase) were also upregulated, particularly with AsA. These treatments improved growth, yield traits, and water use efficiency, especially under drought stress, with Nubaria-5 showing the highest drought resilience. This cultivar exhibited superior pigment stability, antioxidant activity, and yield preservation across stress conditions. Significant interactions among irrigation regime, biostimulant, and cultivar highlighted the importance of genotype-specific responses. Heatmap analysis confirmed the consistent effectiveness of AsA, particularly under severe drought in Nubaria-5. Overall, AsA and HA function as effective biostimulants for enhancing drought resilience in faba bean by improving photosynthetic efficiency, water relations, and antioxidative capacity, with AsA showing greater overall efficacy.
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spelling doaj-art-075e4d61c99b4bfa8049e7999929c9e42025-08-20T04:01:53ZengBMCBMC Plant Biology1471-22292025-08-0125111910.1186/s12870-025-06971-3Enhancing drought tolerance in faba bean using ascorbic and humic acids: role of antioxidant enzymes and compatible solutesHussein S. Al-Demrdash0Anam Ayyoub1Omar E. A. ziton2Saber A.E. Mowafy3El-Sayed E. A. El-Sayed4Uthman Balgith Algopishi5Ahmed Ezzat Ahmed6Khaled A. El-Tarabily7Synan F. AbuQamar8Mohsin Mahmood9El-Sayed M. Desoky10Department of Crop Science, Faculty of Agriculture, Zagazig UniversityCollege of Life Sciences, Northwest A&F UniversityDepartment of Crop Science, Faculty of Agriculture, Zagazig UniversityDepartment of Crop Science, Faculty of Agriculture, Zagazig UniversityDepartment of Crop Science, Faculty of Agriculture, Zagazig UniversityDepartment of Biology, College of Science, King Khalid UniversityDepartment of Biology, College of Science, King Khalid UniversityDepartment of Biology, College of Science, United Arab Emirates UniversityDepartment of Biology, College of Science, United Arab Emirates UniversityKey Laboratory of Grassland Ecology, School of Ecology and Environment, Inner Mongolia UniversityBotany Department, Faculty of Agriculture, Zagazig UniversityAbstract Drought is a major environmental stress, particularly in arid regions, where it severely limits faba bean productivity. Foliar-applied ascorbic acid (AsA) and soil-applied humic acid (HA) significantly improved drought tolerance in three faba bean cultivars by enhancing physiological performance and mitigating oxidative damage under moderate (300 mm) and severe (200 mm) drought conditions. Drought stress significantly reduced chlorophyll content (up to -57.5%), relative water content (RWC, -37.9%), and yield traits such as plant height (− 9.6%) and seed yield (− 20.8%), while increasing oxidative stress markers like malondialdehyde (MDA, + 192.8%) and hydrogen peroxide (H₂O₂, + 105.0%). AsA and HA alleviated these effects, improving chlorophyll retention (up to + 33.7%), water status (+ 17.0%), and reducing MDA(− 19.1%) and electrolyte leakage (− 11.5%). Enhanced accumulation of proline (+ 27.4%) and soluble sugars (+ 18.0%) contributed to improved osmotic balance, while antioxidant enzyme activities (superoxide dismutase, catalase, peroxidase, ascorbate peroxidase, and glutathione reductase) were also upregulated, particularly with AsA. These treatments improved growth, yield traits, and water use efficiency, especially under drought stress, with Nubaria-5 showing the highest drought resilience. This cultivar exhibited superior pigment stability, antioxidant activity, and yield preservation across stress conditions. Significant interactions among irrigation regime, biostimulant, and cultivar highlighted the importance of genotype-specific responses. Heatmap analysis confirmed the consistent effectiveness of AsA, particularly under severe drought in Nubaria-5. Overall, AsA and HA function as effective biostimulants for enhancing drought resilience in faba bean by improving photosynthetic efficiency, water relations, and antioxidative capacity, with AsA showing greater overall efficacy.https://doi.org/10.1186/s12870-025-06971-3Abiotic stress toleranceOxidative stressWater use efficiencyCultivar toleranceReactive oxygen speciesPhysio-biochemical traits
spellingShingle Hussein S. Al-Demrdash
Anam Ayyoub
Omar E. A. ziton
Saber A.E. Mowafy
El-Sayed E. A. El-Sayed
Uthman Balgith Algopishi
Ahmed Ezzat Ahmed
Khaled A. El-Tarabily
Synan F. AbuQamar
Mohsin Mahmood
El-Sayed M. Desoky
Enhancing drought tolerance in faba bean using ascorbic and humic acids: role of antioxidant enzymes and compatible solutes
BMC Plant Biology
Abiotic stress tolerance
Oxidative stress
Water use efficiency
Cultivar tolerance
Reactive oxygen species
Physio-biochemical traits
title Enhancing drought tolerance in faba bean using ascorbic and humic acids: role of antioxidant enzymes and compatible solutes
title_full Enhancing drought tolerance in faba bean using ascorbic and humic acids: role of antioxidant enzymes and compatible solutes
title_fullStr Enhancing drought tolerance in faba bean using ascorbic and humic acids: role of antioxidant enzymes and compatible solutes
title_full_unstemmed Enhancing drought tolerance in faba bean using ascorbic and humic acids: role of antioxidant enzymes and compatible solutes
title_short Enhancing drought tolerance in faba bean using ascorbic and humic acids: role of antioxidant enzymes and compatible solutes
title_sort enhancing drought tolerance in faba bean using ascorbic and humic acids role of antioxidant enzymes and compatible solutes
topic Abiotic stress tolerance
Oxidative stress
Water use efficiency
Cultivar tolerance
Reactive oxygen species
Physio-biochemical traits
url https://doi.org/10.1186/s12870-025-06971-3
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